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The mailbox that is forwarding and the mailboxes it's forwarding to were migrated from Exchange 2003 to 2010.

Maybe the attributes are actually set but are not showing correctly in EMC for some reason?

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Have you checked for any rules in the email client? –  joeqwerty Jun 10 '11 at 16:53

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Are you sure you don't have the "forwarding" mailbox's email address added as an email alias on the receiving mailbox?

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